In this world of Pokémon, there have lived two Guardians that had served under Arceus to protect the world and teach humans and Pokémon how to live side by side together. Now in present time, the Guardians have been reincarnated and begin to learn of their destiny. Come along with our heroes as they discover their powers and meet many Pokémon that thrive in this world!

It has been a few months since the invasion and attack of Calamity Tabuu at the Mansion, causing the Smashers to turn into Trophies and being sent to parts of their worlds, except for one survivor. With her Trophy Slate in hand, join Taylor as she sets off to find every fighter and prepare for the final showdown against their true enemy!