Finding himself on the run for stealing (again), Awful Alvin then finds himself under the protection of a young street teen named Mira. But this girl has secrets, and these secrets could change both their lives forever.

Feeling homesick for Madagascar, King Julien decides to return to Madagascar along with his lemurs, the penguins, and Marlene. However, what was supposed to be nice vacation quickly takes a turn when numerous secrets from the past and present come to light. Especially where Marlene is involved...