Lily is an odd Inkling who's learning to live in the big city of Inkopolis among other marine life. She's also ex-human, flung far into the future from her own time. But it's hard to keep a secret like that from the New Squidbeak Splatoon, who found her; Off the Hook, who think she's connected to the NILS facility; and DJ Octavio, who wants her and her history for himself.