Ultraman Dyna/Shin Asuka crosses into yet another dimension, and ends up befriending a young teen named Shinji Ikari. As the mysterious aliens known as Angels arise and the defense forces of this new Earth deploy the cyborg, Evangelion, Shinji finds himself inheriting the light of his own world's Ultraman.

Ten years since the defeat of Dark Axis, Neotopia and the rest of the dimensions are at peace. However, a new threat of darkness rises up to destroy the Gundam Force, which has recovered an unknown artifact of great power. In the wings, an amnesiac white robot with a single horn is about to discover his destiny as the dimensions barrel towards war once more.